Fish Eating Birds and Salmonids in Scotland

REFERENCES

Alcorn, J.R. 1953. Food of the common merganser in Churchill County, Nevada. Condor 55:151-152.

Alexander, G.D. 1995. Foraging ecology of goosander (Mergus merganser) and red-breasted merganser (M. serrator) ducklings in North east Scotland. PhD Thesis, University of Aberdeen, pp126.

Allison, A., Newton, I. & Campbell, C. 1974. Loch Leven NNR, a study of waterfowl biology. Wildfowlers Association of Great Britain, Chester.

Andrews, I. 1987. Musselburgh Ash Lagoons. Scott. Bird News 7:12-13.

Anderson, J.M. 1986. Merganser predation and its impact on Atlantic salmon stocks in the Restigouche river system 1982-1985. Atlantic Salmon Federation, Special Publication Series No 13, September 1986.

Armstrong, J.D. 1997. Self-thinning in juvenile sea trout and other salmonid fishes revisited. Journal of Animal Ecology 66, 519-526.

Armstrong, J.D., Shackley, P.E. & Gardiner, R. 1994. Redistribution of juvenile salmonid fishes after localized catastrophic depletion. Journal of Fish Biology 45, 1027-1039.

Armstrong, J.D. & Stewart, D.C. 1996. The relationship between first vertebra width and body length of Atlantic salmon differs between parr and smolts. Journal of Fish Biology 49:1038-1040.

Armstrong, J.D., Braithwaite, V.A. & Rycroft, P. 1996. A flat-bed passive integrated transponder array for monitoring behaviour of Atlantic salmon parr and other fish. Journal of Fish Biology 48, 539-541.

Armstrong, J.D. & Stewart, D.C. 1997. The effects of initial length and body curvature on shrinkage of juvenile Atlantic salmon during freezing. Journal of Fish Biology 50: 903-905.

Armstrong, J.D., Braithwaite, V.A., and Huntingford, F.A. 1997. Spatial strategies of wild Atlantic salmon parr: exploration and settlement in unfamiliar areas. Journal of Animal Ecology 66, 203-211.

Armstrong, J.D., Braithwaite, V.A. & Fox, M. 1998. The response of wild Atlantic salmon parr to acute reductions in water flow. Journal of Animal Ecology 67, 292-297.

Aspinall, S.J. & Dennis, R. 1988. Goosander and red-breasted mergansers in the Moray Firth. Scottish Birds 15:65-70.

Atkinson, K.M. & Hewitt, D.P. 1978. A note on the food consumption of red-breasted merganser. Wildfowl 29:87-91.

Bagge, P., Lemmetyinen, R. & Raitis, T. 1973. Spring food of some diving waterfowl in the southwestern Finnish archipelago. Oikos 15:146-150.

Barlow, C.G. & Bock, K. 1984. Predation of fish in farm dams by cormorants Phalacrocorax spp. Australian Wildlife Research 11:559-566.

Barrett, R.T., Rov, N. Loen, J.& Montevecchi, W.A. 1990. Diets of shags Phalacrocorax. aristotalis and cormorants P. carbo in Norway and possible implications for gadoid stock recruitment. Marine Ecology - Progress Series 66:205-218.

Beard, T.D. & Carline, R.F. 1991. Influence of spawning and other stream habitat features on spatial variability of wild brown trout. Transactions of the American Fisheries Society 120, 711-722.

Begon, M., Harper, J.L. & Townsend, C.R. 1996. Ecology: individuals, populations and communities. 3rd Edition. Blackwell Science, Oxford.

Bengtson, S-A. 1971. Food and feeding of diving ducks breeding at lake Myvatn, Iceland. Ornis Fennica 48:77-92.

Bengtson, S-A. 1972. Reproduction and fluctuations in the size of duck populations at lake Myvatn, Iceland. Oikos 23:35-58.

Bergman, G. 1956. Om kullsammanslagning hos skrakar Mergus serrator och Mergus merganser. Fauna Och Flora 3:97-110.

Berry, J. 1936. British mammals and birds as enemies of the Atlantic salmon Salmo salar. Avon Biological Research, Annual Report 1934-35 :31-64.

Berry, J. 1939. The status and distribution of wild geese and wild duck in Scotland. Cambridge University Press.

Bianki, V., Boiko, N. & Kokhanov, V. 1997. The cormorant Phalacrocorax carbo in Kandalaksha Bayn (White Sea, Russia). Ekologia Polsks 45(1): 15.

Bjornn, T.C. 1971. Trout and salmon movements in two Idaho streams as related to temperature, food, stream flow, cover, and population density. Transactions of the American Fisheries Society 100, 423-438.

Bjornn, T.C. & Kirking, S.C. 1991. Relation of cover alterations to the summer standing crop of young salmonids in small southeast Alaska streams. Transactions of the American Fisheries Society 120, 562-570.

Boer, H. de 1972. De voedselbiologie van aalscholver. Rijksinstituut voor Natuurbeheer, Leersum.

Bohlin, T., Dellefors, C., Faremo, U. & Johlander, A. 1994. The energetic equivalence hypothesis and the relation between population density and body size in stream-living salmonids. The American Naturalist 143, 478-493.

Boldreghini, P., Santolini, R. & Pandolfi, M. 1997. Abundance and frequency of occurrence of fish-prey in the diet of cormorants Phalacrocorax carbo in the Po River Delta (Northern Italy) during the wintering period. Ekologia Polska 45(1):191-196.

Boyd, H. 1959. Movements of marked sea and diving ducks in Europe. Wildfowl Trust 10th Annual Report 59-70.

Buck, R.J.G. and Hay, D. 1984. The relationship between stock size and progeny of Atlantic salmon, Salmo salar L., in a Scottish stream. Journal of Fish Biology 24, 1-12.

Buck, R.J.G. and Youngson, A.F. 1982. The downstream migration of precociously mature Atlantic salmon, Salmo salar L., parr in autumn: its relation to the spawning migration of mature adult fish. Journal of Fish Biology 20, 279-288.

Carss, D.N. 1989. Sawbill ducks at fish farms in Argyll, western Scotland. Scottish Birds 15:145-150.

Carss, D.N. 1990. 'Beak prints' help in war against aerial invaders. Fish Farmer 13:46-47.

Carss, D.N. 1993a. Shags Phalacrocorax aristotelis at cage fish farms in Argyll, western Scotland. Bird Study 40:203-211.

Carss, D.N. 1993b. Cormorants Phalacrocorax carbo at cage fish farms in Argyll, western Scotland. Seabird 15:38-44.

Carss, D.N. 1993c. Grey heron Ardea cinerea L. predation at cage fish farms in Argyll, western Scotland. Aquaculture and Fisheries Management 24:29-45.

Carss, D. N 1994. Killing of piscivorous birds at Scottish fin fish farms 1984-7. Biological Conservation 68:181-188.

Carss, D.N.& Brockie, K. 1994. Prey remains at Osprey nests in Tayside and Grampian, 1987-1993. Scottish Birds 17:132-145.

Carss, D.N.& Elston, D.A. 1996. Errors associated with otter Lutra lutra faecal analysis. II. Estimating prey size distribution from bones recovered in spraints. Journal of Zoology, London 238:319-332.

Carss, D.N. & Marquiss, M. 1992. Avian predation at farmed and natural fisheries. Proc. 22nd Annual Institute of Fisheries Management Study Course. Aberdeen 1991.

Carss, D.N. & Marquiss, M. 1996a. The influence of a fish farm on grey heron Ardea cinerea breeding performance. in Aquatic Predators and their Prey (Eds S.P.R.Greenstreet & M.L.Tasker) Blackwell Scientific Publications. Oxford.

Carss, D.N. & Marquiss, M. 1996b. Fish-eating birds: perceptions and 'realities'. Proc. 1995 British Trout Farming Conference. Sparsholt College, Hampshire, Sept. 1995.

Carss, D.N., Bevan, R.M., Bonetti, A., Cherubini, G., Davies, J., Doherty, D., El Hili, A., Feltham, M.J., Grade, N., Granadeiro, J.P., Gremillet, D., Gromadzka, J., Harari, Y., Holden, T., Keller, T., Lariccia, G., Mantovani, R., McCarthy, T.M., Mellin, M., Menke, T., Mirowska-Ibron, I., Muller, W., Musil, P., Nazirides, T., Suter, W., Trauttmansdorff, J.F.G., Volponi, S. & Wilson, B. (1997a) Techniques for assessing cormorant diet and food intake: towards a consensus view. Supplemento Ricerche di Biologia della Selvaggina 26:197-230.

Carss, D.N. & Marquiss, M. 1997. The diet of cormorants Phalacrocorax carbo in Scottish freshwaters in relation to feeding habitats and fisheries. Ekologia Polska 45(1):207-222.

Carss, D.N., Marquiss, M. & Lauder, A. 1997b. Cormorant Phalacrocorax carbo carbo predation at a major trout fishery in Scotland. Proc. Supplemento Ricerche di Biologia della Selvaggina 26:281-294.

Carter, S.P. 1993. Sawbills in Britain in 1987. In: Andrews, J. & Carter, S.J. (Eds) Britain's Birds 1990/91. JNCC/BTO. Peterborough.

Carter, S.P. & Evans, P.R. 1986. Goosander and merganser studies in Scotland. Final report on contract HP3-03-208/9. Nature Conservancy Council. Peterborough.

Chandler, R.J. 1981. Influxes into Britain and Ireland of red-necked grebes and other waterbirds during winter 1978/79. British Birds 74:55-81.

Cherrett, J.M. 1989. Ecological concepts. Blackwell Scientific Publications, London.

Coldwell, C. 1939. The feeding habits of American mergansers. Canadian Field Naturalist 53:55.

Cooper, J. 1986. Diving patterns of cormorants Phalacrocoridae. Ibis 128:562-570.

Cosgrove, P.J. 1997. A winter survey of sawbill ducks and cormorants on the River Deveron, north east Scotland. Scottish Birds 19:93-100.

Cowx, I.G. and Lamarque, P. 1990. Fishing with Electricity. Oxford: Fishing News Books, Blackwell Scientific Publications, Ltd.

Cramp, S. & Simmons, K.E.L. 1977. Handbook of the birds of Europe, the Middle East and North Africa Vol. I. Oxford University Press, Oxford.

Crawley, M.J. 1992. Natural enemies: the population biology of predators, parasites and diseases. Blackwell Scientific Publications, London.

Crozier, W.W & Kennedy, G.J.A. 1993. Marine survival of wild and hatchery-reared Atlantic salmon (Salmo salar L.) from the River Bush, Northern Ireland. In Salmon in the Sea and New Enhancement Strategies (Ed D.H. Mills) Fishing News Books, Oxford.

Dement'ev G.P. & Gladkov, N.A. (Eds) 1952. Birds of the Soviet Union Vol IV. Sovetskaya Nauka, Moskva.

Dirksen, S., Boudewijn, T.J., Noordhuis, R & Marteijn, E.C.L. 1995. Cormorants Phalacrocorax carbo in shallow eutrophic freshwater lakes: prey choice and fish consumption in the non-breeding period and effects of large-scale fish removal. Area 83(1):167-184.

Dickson, R.C. 1987. Red-breasted mergansers moulting in Galloway. Scottish Birds 14:214.

Dill, L.M. 1987. Animal decision making and its ecological consequences: the future of aquatic ecology and behaviour. Canadian Journal of Zoology 65:803-811.

van Dobben, W.H. 1952. The food of the cormorant in the Netherlands. Ardea 40:1-63.

Doornbos, G. 1984. Piscivorous birds on the saline lake Grevelingen, The Netherlands: abundance, prey selection and annual food consumption. Netherlands Journal of Sea Research 18:457-479.

Duffy, D.C. & Jackson, S. 1986. Diet studies of seabirds: a review of methods. Colonial Waterbirds 9:1-17.

Egglishaw, H.J. & Shackley, P.E. 1977. Growth, survival and production of juvenile salmon and trout in a Scottish stream, 1966-75. Journal of Fish Biology 11, 647-672.

Egglishaw, H.J. & Shackley, P.E. 1982. Influence of water depth on dispersion of juvenile salmonids, Salmo salar L. and S. trutta L., in a Scottish stream. Journal of Fish Biology 21, 141-155.

Elliott, J.M. 1986. Spatial distribution and behavioural movements of migratory trout Salmo trutta in a Lake District stream. Journal of Animal Ecology 55, 907-922.

Elliott, J.M. 1993. The self-thinning rule applied to juvenile sea trout, Salmo trutta. Journal of Animal Ecology 62, 371-379.

Elliott, J.M. 1994. Quantitative Ecology and the Brown Trout. Oxford University Press, Oxford.

Englert, J. & Seghers, B.H. 1983. Predation by fish and common mergansers on darters (Pisces: Percidae) in the Thames watershed of Southwest Ontario. Canadian Field Naturalist 97:218-219.

Elson, P.F. 1962. Predator-prey relationships between fish-eating birds and Atlantic salmon. Bulletin of the Fisheries Research Board of Canada 133:87pp.

Erkinaro, J. 1995. Growth differences between the Atlantic salmon parr, Salmo salar, of nursery brooks and natal rivers in the River Teno watercourse in northern Finland. Environmental Biology of Fishes 42, 277-287.

Erkinaro, H. and Erkinaro, J. 1998. Feeding of Atlantic salmon, Salmo salar L., parr in the subarctic River Teno and three tributaries in northernmost Finland. Ecology of Freshwater Fish 7, 13-24.

Erskine, A.J. 1971. Growth and annual cycles and weights, plumages and reproductive organs, of goosanders in eastern Canada. Ibis 113:42-58.

Erskine, A.J. 1972. Populations, movements and seasonal distribution of mergansers. Canadian Wildlife Service Report Series No.17:1-35.

Evans, R.J. 1998. Numbers of wintering seaducks, divers and grebes in the Moray Firth, 1977-1995. Scottish Birds 19:206-222.

Fausch, K.D. 1984. Profitable stream positions for salmonids: relating specific growth rate to net energy gain. Canadian Journal of Zoology 62, 441-451.

Feltham, M.J. 1990. The diet of red-breasted mergansers Mergus serrator during the smolt run in northeast Scotland: the importance of salmon salmo salar smolts and parr. Journal of Zoology 222:285-292.

Feltham, M.J. 1995a. Consumption of Atlantic salmon smolts and parr by goosanders: estimates from doubley-labelled water measurements of captive birds released on two Scottish rivers. Journal of Fish Biology 46:273-281.

Feltham, M J. 1995b. Predation of Atlantic salmon, Salmo salar L., smolts and parr by red-breasted mergansers, Mergus serrator L., on two Scottish rivers. Fisheries Management and Ecology. 2:289-298.

Feltham, M.J. & Davies, J.M. 1997. Daily food intake of cormorants: a summary. Supplemento Ricerche di Biologia della Selvaggina 26:259-268.

Feltham, M J. & MacLean, J.C. 1996. Carlin tag recoveries as an indicator of predation on salmon smolts by goosanders and red-breasted mergansers. Journal of Fish Biology 48:270-282.

Feltham, M.J. & Marquiss, M. 1989. The use of first vertebrae in separating and estimating the size of, trout (Salmo trutta) and salmon (Salmo salar) in bone remains. Journal of Zoology 219:113-122.

Fritsch, L.E. & Bus, I.O. 1958. Food of the American merganser in Unakwik Inlet, Alaska.Condor 60:410-411.

Garcia de Leaniz, C. 1989. Site fidelity and homing of Atlantic salmon parr in a small Scottish Stream. In: E. Brannon & B. Jonsson (eds.). Salmonid Migration and Distribution Symposium. University of Washington Press, Seattle, 70-80.

Gardarsson, A 1979. Waterfowl populations of lake Myvatn and recent changes in numbers and food habits. Oikos 32:250-270.

Gardiner, W.R. 1983. Estimating population densities of salmonids in deep water in streams. Journal of Fish Biology 24, 41-49.

Gardiner, R. and Shackley, P. 1991. Stock and recruitment and density-dependent growth of salmon, Salmo salar L., in a Scottish stream. Journal of Fish Biology 38, 691-696.

Geroudet, P. 1985. Essai de synthese sur l'evolution du harle bievre Mergus merganser dans le bassin du Leman. Nos Oiseaux 38:1-18.

Gibbons, D.J., Reid, J.B. & Chapman, R.A. 1993. The New Atlas of Breeding Birds in Britain and Ireland: 1988-91. Academic Press.

Gibson, R.J. 1993. The Atlantic salmon in fresh water. Spawning, rearing and production. Reviews in Fish Biology and Fisheries 3, 39-73.

Gowan, C. & Fausch, K.D. 1996. Mobile brook trout in two high elevation Colorado streams: re-evaluating the concept of restricted movement. Canadian Journal of Fisheries and Aquatic Sciences 53, 1370-1381.

Gowan, C., Young, M.K., Fausch, K.D. & Riley, S.C. 1994. Restricted movements in residential stream salmonids: a paradigm lost? Canadian Journal of Fisheries and Aquatic Sciences 51, 2626-2637.

Grant, J.W.A. 1993. Self-thinning in stream-swelling salmonids, p. 99-102. In R.J. Gibson and R.E. Cutting (ed.) Production of juvenile Atlantic salmon, Salmo salar, in natural waters. Canadian Special Publication in Fisheries and Aquatic Sciences 118.

Grant, J.W.A., Steingrimsson, S.Ó., Keeley, E.R. & Cunjak, R.A. 1998. Territory size and its implications for the measurement and prediction of salmonid abundance. Canadian Journal of Fisheries and Aquatic Sciences. In press.

Grant, J.W.A. & Kramer, D.L. 1990. Territory size as a predictor of the upper limit to population density of juvenile salmonids in streams. Canadian Journal of Fisheries and Aquatic Sciences 47, 1724-1737.

Gregory, R.D., Carter, S.P. & Baillie, S.R. 1997. Abundance,distribution and habitat use of breeding goosanders Mergus merganser and red-breasted mergansers Mergus serrator on British rivers. Bird study 44:1-12.

Gremillet, D., Schmid, D. & Culik, B. 1995. Energy requirements of breeding great cormorants Phalacrocorax carbo sinensis. Marine Ecology - Progress Series 121:1-9.

Griffin, B. 1990. Breeding sawbills on Welsh rivers 1990. RSPB report, 15pp.

Gustafson-Greenwood, K.I. & Moring, J.R. 1990. Territory size and distribution of newly-emerged Atlantic salmon (Salmo salar). Hydrobiologia 206, 125-131.

Hansen, S.G. 1980. Breeding status of the goosander in Norway. Danske Fugle 32:147-151.

Hansen, L.P, Jonsson, B. & Jonsson, N. 1996. Atlantic salmon of the rivers Imsa and Drammen. NINA Oppdragsmelding 401, 1-28.

Harcup, M.F., Williams, R. & Ellis, D.M. 1984. Movements of brown trout, Salmo trutta L., in the River Gwyddon, South Wales. Journal of Fish Biology 24, 415-426.

Harkonen, T.J. 1988. Food-habitat relationship of harbour seals and black cormorants in Skagerrak and Kattegat. Journal of Zoology 214:673-681.

Harris, M.P. & Wanless, S. 1993. The diet of Shags Phalacrocorax aristotalis during the chick-rearing period assessed by three methods. Bird Study 40:135-139.

Hartley, P.H.T. 1948. The assessment of the food of birds. Ibis 90:361-381.

Heard, W.R. & Curd, M.R. 1959. Stomach contents of American mergansers Mergus merganser caught in gill nets set in Lake Carl Blackwell, Oklahoma. Proceedings - Oklahoma Academy of Science 39:197-200.

Heggenes, J. 1988. Effect of experimentally increased intraspecific competition on sedentary adult brown trout (Salmo trutta) movement and stream habitat choice. Canadian Journal of Fisheries and Aquatic Sciences 45, 1163-1172.

Heggenes, J., Northcote, T.G. & Peter, A. 1991. Spatial stability of cutthroat trout (Oncorhynchus clarki) in a small coastal stream. Canadian Journal of Fisheries and Aquatic Sciences 48, 757-762.

Hesthagen, T. 1988. Movements of brown trout, Salmo trutta, and juvenile Atlantic salmon, Salmo salar, in a coastal stream in northern Norway. Journal of Fish Biology 32, 639-653.

Heyer, R.A., Donnelly, M.A., McDiarmid, R.W., Hayek, L-A, C. & Foster, M.S. 1994. Measuring and monitoring biological diversity: standard methods for amphibians. Smithsonian Institution Press, London.

Hilden, O. 1964. Ecology of duck populations in the island group of Valassaaret, Gulf of Bothnia. Annales Zoologici Fennica 1:153-227.

Hislop, J.R.G. & MacDonald, W.S. 1989. Damage to fish by seabirds in the Moray Firth. Scottish Birds, 15:151-155.

Hofer, J. & Marti, C. 1988. Beringungsdaten zur uberwinterung des gansesagers Mergus merganser am Sempachersee: herkunft, zugverhalten und gerwicht. Ornithologische Beobachter 85:97-122.

Huntingdon, E.H. & Roberts, A.A. 1959. Food habits of the merganser in New Mexico. Bull. No. 9. New Mexico dept. Game & Fish. Santa Fe.

Huntingford, F.A., Braithwaite, V.A., Armstrong, J.D., Aird, D., Thorpe, K.E. & Joiner, P. 1998. Social status and growth rates as determinants of site attachment in juvenile Atlantic salmon. Journal of Fish Biology 53:314-321.

Im, B.H. & Hafner, H. 1984. Impact des Oiseaux piscivores et plus particulierement du grand cormoran Phalacrocorax carbo sinensis sur les exploitations piscicoles en Camargue, France. Le Sambuc, Station Biologique de la Tour du Valat.

Jenkins, D.J. (Ed) 1985. The biology and management of the River Dee. ITE Symposium 14. ITE, Huntingdon.

Jenkins, T.M. 1969. Social structure, position choice and micro-distribution of two species (Salmo trutta and Salmo gairdneri) resident in mountain streams. Animal Behaviour Monographs 2, 57-123.

Kalas, J.A., Heggberget, T.G., Bjorn, P.A. & Reitan, O. 1993. Feeding behaviour and diet of goosanders Mergus merganser in relation to salmonid seaward migration. Aquatic Living Resources 6:31-38.

Keeley, E.R. & Grant, J.W.A. 1995. Allometric and environmental correlates of territory size in juvenile Atlantic salmon (Salmo salar). Canadian Journal of Fisheries and Aquatic Sciences 52, 186-196.

Keller, T. (1997). Food of cormorants Phalacrocorax carbo wintering in Bavaria (southern Germany). Ekologia Polska 45(1):237-238.

Kennedy, G.J.A. & Greer, J.E. 1988. Predation by cormorants Phalacrocorax carbo (L) on the salmonid populations of an Irish river. Aquaculture and Fisheries Management 19:159-170.

Kennedy, G.J.A and Strange, C.D. 1982 The distribution of salmonids in upland streams in relation to depth and gradient. Journal of Fish Biology 20, 579-592.

von Kieckbusch, J.J. 1993. Beobachtungen zur nahrungswahl des kormorans Phalacrocorax carbo sinensis in der umgebung des naturschutzgebietes ' Oehe-Schleimunde'. Seevogel, Zeitschrift Verein Jordsand, Hamburg 1993. 14:19-22.

Kirby, J.S. 1995. Winter population estimates for selected waterfowl species in Great Britain. Biological Conservation 73:189-198.

Kirby, J.S., Salmon, D.G., Atkinson-Willes, G.L. & Cranswick, P.A. 1995. Index numbers for waterbird populations. III. Long-term trends in the abundance of wintering wildfowl in Great Britain, 1966/67-1991/92. Journal of Applied Ecology 32:536-551.

Lack, P. (1986) The atlas of wintering birds in Britain and Ireland. Poyser. Calton.

Leonard, J.W. & Shetter, D.S. 1936. Studies on merganser depredations in Michigan trout waters. Transactions of the American Fisheries Society 66:335-337.

Lid, G. & Schandy, T. 1984. Laksandas forekomst og nearingsvalg I Hallingdalselva ved gol. 30pp NVE-Vassdragsdirectoratet.

Lindroth, A. 1955. Mergansers as salmon and trout predators in the river Indalsalven. Report: Institue of Freshwater Research Drottingholm 36:126-132.

Lindroth, A. & Bergstom, E. 1959. Notes on the feeding technique of the goosander in streams. Fishery Board of Sweden, Report: Institue of Freshwater Research Drottingholm Rept. 40:165-175.

Little, B.& Furness, R.W. 1985. Long distance moult migration by British goosanders. Ringing and Migration 6:214-216.

Macdonald, R.A. 1987a. The breeding population and distribution of the cormorant in Ireland. Irish Birds 3:405-416.

Macdonald, R.A. 1987b. Red-breasted merganser's large creche. Scottish Bird News 6:4.

McIntosh, R. 1978. Distribution and diet of the cormorant on the lower reaches of the river Tweed. Fisheries Management 9:107-113.

Madsen, F.J. 1957. On the food habits of some fish-eating birds in Denmark. Danish Review of Game Biology 3:19-83.

Madsen, F.J. & Sparck, R. 1950. On the feeding habits of the southern cormorant (Phalacrocorax carbo sinensis Shaw) in Denmark. Danish Review of Game Biology 1(3):45-75.

Maitland, P.S. & Campbell, R.N. 1992. Freshwater Fishes of the British Isles. Harper Collins, London.

Marquiss, M. & Carss, D.N. 1994. Avian Piscivores: Basis for Policy. National Rivers Authority Research & Development Report 461/8/N&Y.

Marquiss, M. & Carss, D.N. 1997. Methods of estimating the diet of sawbill ducks Mergus spp. and cormorants Phalacrocorax carbo. Supplemento Ricerche di Biologia della Selvaggina 26:247-258.

Marquiss, M & Duncan, K. 1993. Variation in the abundance of red-breasted mergansers on a Scottish river in relation to season, year, river hydrography, salmon density and spring culling. Ibis 135:33-41.

Marquiss, M & Duncan, K. 1994a. Seasonal switching between habitats and changes in abundance of goosanders Mergus merganser within a Scottish river system. Wildfowl 45:198-208.

Marquiss, M & Duncan, K. 1994b. Diurnal activity patterns of goosanders Mergus merganser on a Scottish river system. Wildfowl 45:209-221.

Marquiss, M., Feltham, M.J. & Duncan, K. 1991. Sawbill ducks and Salmon. Fisheries Research Services Report No. 18/91. Scottish Office Agriculture & Fisheries Department. Aberdeen.

Marquiss, M. & Leitch, A.F. 1990. The diet of grey herons Ardea cinerea breeding at Loch Leven, Scotland, and the importance of their predation on ducklings. Ibis 132:535-549.

Martin, M.W.A. 1988. Colonial nesting red-breasted mergansers. Scottish Bird News 9:8.

Martucci, O. & Giovacchini, P. (1994). Some aspects of the feeding habits of the cormorant Phalacrocorax carbo wintering in Maremma Natural Park (GR, Central Italy). Avocetta 18:53-56.

Martyniak, A., Mellin, M., Stachowiack, P. & Wittke, A. (1993). Food composition of cormorants in two colonies in NE Poland from pellet analysis. Abstracts: Proceedings Workshop 1993 on Cormorants Phalacrocorax carbo. 13-17 April, in Gdansk, Poland.

Meigs, R.C. & Rieck, C.A. 1967. Mergansers and trout in Washington. Proceeding of the 47th Annual Conference Western Association State Game and Fish Commissioners. pp306-318.

Mellin, M. & Martyniak, A. 1991. Food composition of the cormorant Phalacrocorax carbo sinensis in northeastern Poland; preliminary results. Proc. Workshop 1989 on Cormorants Phalacrocorax carbo. Van Eerden, M.R. & Zijlstra, M. (Eds). Rijkswaterstaat Directorate Flevoland. Lelystad:175-181.

Mellin, M., Ibron-Mirowska, I & Martyniack, A. (1997). Food composition of cormorants Phalacrocorax carbo shot at two fish-farms in North-east Poland. Ekologia Polska 45(1):247.

Meek, E.R. & Little, B. 1977a. Ringing studies of goosanders in Northumberland. British Birds 70:273-283.

Meek, E.R. & Little, B.1977b. The spread of the goosander in Britain and Ireland. British Birds 70:229-237.

Meek, E.R. & Little, B. 1980. Goosander studies in the British Isles. Danske Fugle 32:132-146.

Milinski, M. 1986. Constraints placed by predators on feeding behaviour. pp 236-252 in The behaviour of teleost fishes. Ed T.J.Pitcher. Croom Helm Ltd., London.

Miller, S.W. & Barclay, J.S. 1973. Predation in warm water reservoirs by wintering common mergansers. Proceeding of the 27th Annual Conference South Eastern Association State Game Fish Commissioners pp243-252.

Mills, D.H. 1962a. The goosander and red-breasted merganser as predators of salmon in Scottish waters. Freshater and Salmon Fisheries Research 29. HMSO, Edinburgh.

Mills, D.H. 1962b. The goosander and red-breasted merganser in Scotland. Wildfowl Trust 13th Report 79-92.

Mills, D.H. 1965. The distribution and food of the cormorant Phalacrocorax carbo in Scottish inland waters. Freshwater and Salmon Fisheries Research 35. HMSO Edinburgh.

Mills, D.H. 1969. The food of the cormorant at two breeding colonies on the east coast of Scotland. Scottish Birds 5:272-276.

Mittelbach, G.G. 1986. Predation mediated habitat use: some consequences for species interactions. Environmental Biology of Fishes 16:159-169.

Moerbeek, D.J., van Dobben, W.H., Osieck, E.R., Boere, G.C. & Bungenberg de Jong, C.M. 1987. Cormorant damage prevention at a fish farm in the Netherlands. Biological Conservation 39:23-38.

Monval, J.-Y. & Pirot. J.-Y. 1989. Results of the IWRB International Waterfowl Census 1967-86. IWRB Special Publication 8. International Waterfowl and Wetlands Research Bureau, Slimbridge.

Mortensen, E. 1977. The population dynamics of young trout (Salmo trutta L.) in a Danish brook. Journal of Fish Biology 10, 23-33.

Mott, D.F. & Boyd, F.L. 1995. A review of techniques for preventing cormorant depredations at aquaculture facilities in the Southeastern United States. Colonial Waterbirds 18 (Special publication 1):176-180.

Munro, J.A. & Clemens, W.A. 1937. The American merganser in British Columbia and its relation to the fish population. Bulletin of the Fisheries Research Board Canada No. 55:1-50.

Murray, R.D. 1988. The 1987 sawbill and river birds census. in 'The 1987 Borders bird report', Ed R.D.Murray. Scottish Ornithologists Club, Edinburgh.

Murray, R.D. 1992. The 1991 goosander counts. in 'The Borders bird report No. 13.' Ed R.D.Murray, Scottish Ornithologists Club, Edinburgh.

Nilsson, L. 1974. Food choice of resting and wintering goosanders in Scania. Var Fagelvarld 33:293-294

Noordhuis, R., Marteijn, E.C.L. & Noordhuis, R., Dirksen, S. & Boudewijn, T.J. 1997. The trophic role of cormorants Phalacrocorax carbo in freshwater ecosystems in The netherlands during the non-breeding period. Ekologia polska 45(1):249-262.

Niittyla, J. 1980. On the development of some goosander Mergus merganser populations in Finland, a review. Danske Fugle 32:158-165.

Oberdorff, T., Hugueny, B., Compin, A. & Belkessam, D. 1998. Non-interactive fish communities in the coastal streams of North-western France. Journal of Animal Ecology 67:472-484.

Owen, M., Atkinson-Willes, G.L. & Salmon, D.G. 1986. Wildfowl in Great Britain. Cambridge University Press.

Page, L.M. & Burr, B.M. 1991. A field guide to freshwater fishes. Houghton Mifflin Co., Boston, USA.

Parslow, J. 1973. Breeding Birds of Britain and Ireland, a historical survey. Poyser, Berkhamsted.

Peake, S. & McKinley, R.S. 1998. A re-evaluation of swimming performance in juvenile salmonids relative to downstream migration. Canadian Journal of Fisheries and Aquatic Sciences 55:682-687.

Pearson, T.H. 1968. The feeding biology of seabirds breeding on the Farne Islands, Northumberland. Journal of Animal Ecology 37:521-552.

Piggins, D.J. 1959. Investigations on the predators of salmon smolts and parr. Annual Report. Salmon Research Trust Ireland Incorporated 1958. Appendix I, 12pp.

Pitcher, T.J. 1986. Behaviour of teleost fishes. Chapman & Hall, London.

Rad, O. 1980. Breeding distribution and habitat selection of red-breasted merganser in freshwater in western Norway. Wildfowl 31:53-56.

Rae, B.B. 1969. The food of cormorants and shags in Scottish estuaries and coastal waters. Department of Agriculture and Fisheries for Scotland. Marine Research 1. HMSO. Edinburgh.

Rae, S. & Duncan, K. 1989. Goosanders feeding on carrion. Scottish Birds 15:134.

Rehfeldt, G. 1986. Ecology and breeding biology of an internal population of the red-breasted merganser Mergus serrator. Okologie der Vogel 8:133-144.

Reimchen, T.E. 1991. Evolutionary attributes of headfirst prey manipulation and swallowing in piscivores. Canadian Journal of Zoology 69:2912-2916.

Rimmer, D.M., Paim, U. & Saunders, R.L. 1983. Autumnal habitat shift of juvenile Atlantic salmon (Salmo salar) in a small river. Canadian Journal of Fisheries and Aquatic Sciences 40, 671-680.

Rimmer, D.M., Paim, U. & Saunders, R.L. 1984. Changes in the selection of microhabitat by juvenile Atlantic salmon (Salmo salar) at the summer-autumn transition in a small river. Canadian Journal of Fisheries and Aquatic Sciences 41, 469-475.

Robins, C.R., Ray, G.C. & Douglass, J. 1986. Atlantic coast fishes. Houghton Mifflin Co., Boston, USA.

Rose, P.M. & Scott, D.A. 1994. Waterfowl population estimates. IWRB Special Publication 29. International Waterfowl and Wetlands Research Bureau, Slimbridge.

Ruger, A., Prentice, C. & Owen, M. 1986. Results of the IWRB International Waterfowl Census 1967-83. IWRB Special Publication 6. 118pp. International Waterfowl and Wetlands Research Bureau, Slimbridge.

Russell, I.C., Dare, P.J., Eaton, D.R. & Armstrong. 1996. Assessment of the problem of fish-eating birds in inland fisheries in England & Wales. Report to the Ministry of Agriculture, Fisheries and Food (MAFF project VC0104). MAFF, Lowestoft.

St John, C. 1882. Natural History and Sports in Moray. David Douglas. Edinburgh.

Salyer, J.C. & Lagler, K.F. 1940. Food and habits of the American merganser during winter in Michigan , considered in relation to fish management. Journal of Wildlife Management 4:186-219.

Sharrock, J.T.R. 1976. The atlas of breeding birds in Britain and Ireland. BTO. Tring.

Shearer, W.M., Cook, R.M., Dunkley, D.A., Maclean, J.C. & Shelton, R.G.J. 1987. A model to assess the effect of predation by sawbill ducks on the salmon stock of the river North Esk. Scottish Fisheries Research Report 37. HMSO Edinburgh.

Sitters, H.P. 1988. Tetrad atlas of the breeding birds of Devon. Devon Bird Watching Soc. Devon.

Sjoberg, K. 1985. Foraging activity patterns in the goosander Mergus merganser and the red-breasted merganser M. serrator in relation to patterns of activity in their major prey species. Oecologia 67:35-39.

Sjoberg, K. 1988. Food selection, food seeking patterns and hunting success of captive goosanders Mergus merganser and red-breasted mergansers M. serrator in relation to the behaviour of their prey. Ibis 130:79-93.

Sjoberg, K. 1989. Time-related predator/prey interactions between birds and fish in a northern Swedish river. Oecologia 80:1-10.

Snook, C. 1992. Cormorant's method of dealing with large flatfish. British Birds 85:238.

Solomon, D.J. 1982. Migration and dispersion of juvenile brown and sea trout. In: Salmon and Trout Migratory Behavior Symposium. (E.L.Brannon and E.O.Salo, eds). pp. 136-145. University of Washington, Seattle, Washington.

Starling, A.E., Kirby, J.S., Pickering, S.P.C., Gilburn, A.S. & Howell, R. 1992. British status and distribution of wintering cormorants, goosanders and red-breasted mergansers in GreatBritain with specific reference to the Welsh region. National Rivers Authority. Cardiff.

Steven, G.A. 1933. The food consumed by shags and cormorants around the shores of Cornwall (England). Journal of the Marine Biological Association UK 19:227-292.

Stuart, T.A. 1957. The migrations and homing behaviour of brown trout (Salmo trutta L.). Freshwater and Salmon Fisheries Research 18, 1-27

Stutz, S.S. 1965. Size of common merganser broods. The Murrelet. 46:47-48.

Suter, W. 1991. Food and feeding of cormorants Phalacrocorax carbo wintering in Switzerland. Proc.Workshop 1989 on Cormorants Phalacrocorax carbo. Van Eerden, M.R. & Zijlstra, M. (Eds). Rijkswaterstaat Directorate Flevoland. Lelystad:156-165.

Suter, W. 1995. The effect of predation by wintering cormorants Phalacrocorax carbo on grayling Thymallus thymallus and trout (Salmonidae) populations: two case studies from Swiss rivers. Journal of Applied Ecology 32:29-46.

Suter W. 1997. Roach rules: shoaling fish are a constant factor in the diet of cormorants Phalacrocorax carbo in Switzerland. Ardea 85:9-27.

Symons, P.E.K. 1971. Behavioural adjustment of population density to available food by juvenile Atlantic salmon. Journal of Animal Ecology 40, 569-587.

Symons, P.E.K. & Heland, M. 1978. Stream habitats and behavioural interactions of underyearling and yearling Atlantic salmon (Salmo salar). Journal of the Fisheries Research Board of Canada 35, 175-183.

Takashina, H. & Niima,Z. 1957. How the cormorant finds and catches fish and cuts on the fish marked with its beak. Yamashina's Institute of Ornithological Zoology 10:13-17.

Tesch, F.-W. 1977. The eel: biology and management of Anguillid eels. Chapman & Hall. London. pp434.

Timken, R.L. & Anderson, B.W. 1969. Food habits of common mergansers in the northcentral United States. Journal of Wildlife Management 33:87-91.

Tyler, S.J. 1986. Goosanders on the Afon Tywi, the wintering and breeding status of goosanders in wales.RSPB Newtown, Powys.

Tyler, S.J. 1987. Goosanders on the Afon Tywi, monthly counts of goosanders and other piscivores Feb 1986 to Jan 1987. RSPB. Newtown. Powys.

Tyler, S.J., Stratford, J.O. & Lucas, R.M. 1988. Goosanders in Wales. Welsh Bird Report, 1988.

Veldkamp, R. 1995. Diet of cormorants Phalacrocorax carbo sinensis at Wanneperveen, The Netherlands, with special reference to bream Abramis brama. Ardea 83(1):143-155.

Voslamber, B. 1988. Visplaatskeuze, fourageerwijze en voedselkeuze van aalscholvers Phalacrocorax carbo sinensis in het Ijsselmeerebied in 1982. Flevobericht nr. 286. Rijksdienst voor de Ijsselmeerpolders, Lelystad.

West, B., Cabot, D. & Greer-Walker, M. 1975. The food of the cormorant Phalacrocorax carbo at some breeding colonies in Ireland. Proceedings Royal Irish Acadamy 75B:285-304.

Wheeler, A.H. 1978. The Fishes of the British Isles and Northern Europe. Macmillan. London.

White, H.C. 1939. Bird control to increase the Margaree river salmon. Bulletin of the Fisheries Research Board of Canada 58:1-30.

White, H.C. 1957. Food and natural history of mergansers on salmon waters in the Maritime Provinces of Canada. Bulletin of the Fisheries Research Board of Canada 116:1-63.

Wood, C.C. 1985a. Aggregative response of common mergansers Mergus merganser: predicting flock size and abundance on Vancouver Island salmon streams. Canadian Journal of Fisheries and Aquatic Sciences 42:1259-1271.

Wood, C.C. 1985b. Food-searching behaviour of the common merganser Mergus merganser II: Choice of foraging location. Canadian Journal of Zoology 63:1271-1279.

Wood, C.C. 1986. Dispersion of common merganser Mergus merganser breeding pairs in relation to the availability of juvenile Pacific salmon in Vancouver Island streams. Canadian Journal of Zoology 64:756-765.

Wood, C.C. & Hand, C.M. 1985. Food-searching behaviour of the common merganser Mergus merganser I: functional responses to prey and predator density. Canadian Journal of Zoology 63:1260-1270.

Wootton, R.J. 1990. Ecology of teleost fishes. Chapman & Hall, London.

Worthmann, H. & Spratte, S. 1990. Nahrungsuntersuchungen an kormoranen vom Grossen Ploener See. Fischerei Teichwirt. 41:2-8.

Young, D.A. & Titman, R.D. 1988. Intraspecific nest parasitism in red-breasted mergansers. Canadian Journal of Zoology. 66:2454-2458.

Zippin, C. 1958. The removal method of population estimation. Journal of Wildlife Management 22, 82-90.